Definition of ankles:
(n) :
The skeletal joint which connects the foot with the leg; the uppermost portion of the foot and lowermost portion of the leg, which contain this skeletal joint.
(v) :
(slang, ambitransitive) To walk.
(v) :
(cycling) To cyclically angle the foot at the ankle while pedaling, to maximize the amount of work applied to the pedal during each revolution.