Definition of cockles:
(n) :
Any of various edible European bivalve mollusks, of the family Cardiidae, having heart-shaped shells.
(n) :
The shell of such a mollusk.
(n) :
(figurative, in the plural) Chiefly in cockles of someone's heart: a person's innermost feelings.
(n) :
(directly from French coquille) A wrinkle, pucker
(v) :
(transitive) To cause to contract into wrinkles or ridges, as some kinds of cloth after a wetting; to pucker.
(n) :
Any of several field weeds, such as the common corncockle (Agrostemma githago) and darnel ryegrass (Lolium temulentum).